1. Division of Plant Sciences James Hutton Institute University of Dundee School of Life Sciences Errol Rd Invergowrie, Dundee DD2 5DA UK
2. Key Laboratory of Horticultural Plant Biology (HZAU) Ministry of Education Key Laboratory of Potato Biology and Biotechnology (HZAU) Ministry of Agriculture and Rural Affairs Huazhong Agricultural University Wuhan Hubei 430070 China
3. Institute of Molecular, Cell and Systems Biology College of Medical, Veterinary, and Life Sciences University of Glasgow Glasgow G12 8QQ UK
4. Cell and Molecular Science James Hutton Institute Invergowrie, Dundee DD2 5DA UK