1. Bioengineering Sciences Research Group Engineering Sciences Academic Unit Faculty of Engineering and the Environment University of Southampton Southampton SO17 1BJ UK
2. Ecological Sciences Group The James Hutton Institute Invergowrie Dundee DD2 5DA UK
3. Institute of Biological and Environmental Science University of Aberdeen Aberdeen AB24 3UU UK
4. School of Science and Engineering University of Dundee Dundee DD1 4HN UK