1. Department of Marine Sciences; University of North Carolina at Chapel Hill; Chapel Hill NC 27599-3300 USA
2. Land Resources and Environmental Sciences; Montana State University; Bozeman MT 59717 USA
3. Center for Biofilm Engineering; Montana State University; Bozeman MT 59717 USA
4. Faculty of Health, Medicine and Life Sciences; Maastricht University; Maastricht The Netherlands
5. Department of Earth Sciences; University of Southern California; Los Angeles CA 90089 USA
6. Department of Biological Sciences; University of Southern California; Los Angeles CA 90089 USA