1. CNRS, Sorbonne Universités; UPMC Univ Paris 06; Laboratoire d'Océanographie Microbienne (LOMIC); Observatoire Océanologique; F-66650 Banyuls/mer France
2. Graduate program in Oceanography; Department of Oceanography; University of Concepción; Chile
3. Departamento de Biología; Facultad de Ciencias Naturales y Exactas; Universidad de Playa Ancha; Avda. Leopoldo Carvallo 270, Playa Ancha Valparaíso Chile
4. LEMAR-UMR CNRS UBO IRD 6539; Technopole Brest Iroise; Place Nicolas Copernic Plouzané F29280 France
5. Institute for Marine and Antarctic Studies; University of Tasmania; Hobart Tas 7001 Australia
6. Antarctic Climate & Ecosystems Cooperative Research Centre; University of Tasmania; Hobart Tas 7001 Australia