1. School of Civil Engineering and Geosciences; Newcastle University; Newcastle upon Tyne NE1 7RU UK
2. Department of Microbiology & Biogeochemistry, The Royal Netherlands Institute for Sea Research (NIOZ); Den Hag Texel The Netherlands
3. Newcastle University Protein and Proteome Analysis (NUPPA), Devonshire Building, Newcastle University; Newcastle upon Tyne NE1 7RU UK
4. Institute for Geology, Universität Hamburg, Bundesstraße 55; Hamburg 20146 Germany
5. Department of Biology; San Diego State University; San Diego CA USA