1. School of Biomedical and Biomolecular Science; University College Dublin; Belfield; Dublin; 4; Dublin; Ireland
2. Environmental Biology Department; Centro de Investigaciones Biológicas; CSIC; Ramiro de Maeztu 9; 28040; Madrid; Spain
3. School of Biological Sciences and QUESTOR Centre; The Queen's University of Belfast; Medical Biology Centre; 97 Lisburn Road; Belfast; BT9 7BL; UK