1. Institute for Environmental Management Technology; National Institute of Advanced Industrial Science and Technology (AIST); Onogawa 16-1 Tsukuba Ibaraki 305-8569 Japan
2. Bioproduction Research Institute; National Institute of Advanced Industrial Science and Technology (AIST); Higashi 1-1-1 Tsukuba Ibaraki 305-8566 Japan
3. Bioproduction Research Institute; National Institute of Advanced Industrial Science and Technology (AIST); Tsukisamu-higashi 2-17-2-1 Toyohira-ku Sapporo Hokkaido 062-8517 Japan
4. Department of Microbial Ecophysiology; University of Bremen; Leobener Str. NW2 Bremen 28359 Germany