1. Centre for GeoGenetics; Natural History Museum of Denmark; University of Copenhagen; 1350 Copenhagen K Denmark
2. Center for Zoo and Wild Animal Health; Copenhagen Zoo; 2000 Frederiksberg Denmark
3. School of Biological Sciences; University of Bristol; Bristol BS8 1UG UK
4. Trace and Environmental DNA Laboratory; Department of Environment and Agriculture; Curtin University; Perth Western Australia 6102 Australia