1. Institute of Nature Conservation; Polish Academy of Sciences; Al. Mickiewicza 33 31-120 Kraków Poland
2. Evolutionary Biology Group; Faculty of Biology; Adam Mickiewicz University; ul. Umultowska 89 61-614 Poznań Poland
3. Department of Biology; Lund University; Ecology Building, Sölvegatan 37 223 62 Lund Sweden