1. State Key Laboratory of Genetic Resources and Evolution and Yunnan Key Laboratory of Biodiversity and Ecological Security of Gaoligong Mountain Kunming Institute of Zoology, Chinese Academy of Sciences Kunming Yunnan China
2. Kunming College of Life Sciences University of Chinese Academy of Sciences Kunming Yunnan China
3. School of Mathematics and Statistics UNSW Sydney Sydney New South Wales Australia
4. Evolution and Ecology Research Centre, UNSW Sydney Sydney New South Wales Australia
5. Center for Excellence in Animal Evolution and Genetics Chinese Academy of Sciences Kunming Yunnan China
6. School of Biological Sciences University of East Anglia, Norwich Research Park Norwich Norfolk UK