1. Department of Anatomy University of Otago Dunedin New Zealand
2. Department of Zoology University of Otago Dunedin New Zealand
3. Department of Marine Science University of Otago Dunedin New Zealand
4. Trace and Environmental DNA (TrEnD) laboratory, Department of Environment and Agriculture Curtin University Perth WA Australia
5. Department of Biological Sciences Macquarie University North Ryde NSW Australia