1. Department of Evolutionary Biology; Evolutionary Biology Centre (EBC); Uppsala University; Norbyvägen 18D Uppsala 75236 Sweden
2. Department of Biosciences; University of Helsinki; PO Box 65 Helsinki 00014 Finland
3. Centre for Biodiversity Dynamics; Department of Biology; Norwegian University of Science and Technology; Trondheim 7491 Norway
4. Department of Animal Ecology; Evolutionary Biology Centre (EBC); Uppsala University; Norbyvägen 18D Uppsala 75236 Sweden