1. Department of Entomology The University of Arizona Tucson AZ USA
2. Univ Avignon CNRS IRD IMBE Aix Marseille Université Marseille France
3. Department of Zoology University of Oxford Oxford UK
4. BIO5 Institute The University of Arizona Tucson AZ USA
5. Department of Ecology and Evolutionary Biology The University of Arizona Tucson AZ USA