1. Key Laboratory of Exploration and Utilization of Aquatic Genetic Resources Ministry of Education Shanghai Ocean University Shanghai 201306 China
2. Department of Zoology Southern Illinois University Carbondale IL 62901 USA
3. US National Parasite Collection and Animal Parasitic Disease Laboratory USDA Agricultural Research Service BARC East No. 1180, 10300 Baltimore Avenue Beltsville MD 20705 USA
4. Museum of Southwestern Biology and Department of Biology University of New Mexico Albuquerque NM 87131 USA
5. Department of Biology Northern Michigan University Marquette MI 49855 USA