1. Department of Land Resources & Environmental Sciences; Montana State University; P.O. Box 173120 Bozeman MT 59717-3120 USA
2. Institute on Ecosystems; Montana University System; Montana State University; 106 AJM Johnson Hall Bozeman MT 59717 USA
3. Flathead Lake Biological Station; Fish and Wildlife Genomics Group; Division of Biological Sciences; University of Montana; Missoula MT 59812 USA