1. Fisheries Laboratory, Graduate School of Agricultural and Life Sciences University of Tokyo Hamamatsu Japan
2. Department of Applied Aquabiology National Fisheries University Shimonoseki Japan
3. Department of Agriculture, Graduate School of Integrated Arts and Sciences Kochi University Nankoku Japan
4. Okinawa Regional Research Center, Tokai University Taketomi Japan
5. Department of Ecosystem Studies, Graduate School of Agricultural and Life Sciences University of Tokyo Tokyo Japan