1. Department of Biological Sciences; National University of Singapore; 14 Science Drive 4 Singapore 117543 Singapore
2. GeoBioCenter; Ludwig-Maximilians-University; Richard-Wagner-Str. 10 80333 Munich Germany
3. Zoological State Collection; Münchhausenstr 21 D-81247 Munich Germany
4. Evolution, Ecology and Genetics; Research School of Biology; The Australian National University; Canberra ACT 0200 Australia
5. University Scholars Programme; National University of Singapore; 14 Science Drive 4 Singapore 117543 Singapore