1. Key Laboratory of Bio‐Resource and Eco‐Environment of Ministry of Education, College of Life Sciences, State Key Laboratory of Hydraulics and Mountain River Engineering Sichuan University 610065 Chengdu China
2. State Key Laboratory of Grassland Agro‐ecosystem, College of Ecology Lanzhou University 730000 Lanzhou China
3. College of Science Tibet University 850000 Lhasa China
4. College of Biodiversity Conservation Southwest Forestry University 650224 Kunming China