1. The Swire Institute of Marine Science and School of Biological Sciences; The University of Hong Kong; Pokfulam Road Hong Kong SAR China
2. Faculty of Applied Sciences; University of Sunderland; Sunderland SR1 3SD UK
3. Department of Life and Environmental Sciences; Faculty of Science and Technology; Bournemouth University; Fern Barrow Poole BH12 5BB UK
4. Faculty of Biology; University of Vigo; Campus de Vigo 36310 Vigo Spain