1. Department of Ecology and Evolutionary Biology; University of Connecticut; 75 North Eagleville Road Unit 3043 Storrs CT 06269-3043 USA
2. Department of Statistical Science; Duke University; Box 90251 Durham NC 27708-0251 USA
3. Forest Bio-Research Center; Forestry and Forest Products Research Institute; 1 Matsunosato Tsukuba Ibaraki 305-8687 Japan
4. Tama Forest Science Garden; Forestry and Forest Products Research Institute; 1833-81 Todori, Hachioji Tokyo 193-0843 Japan
5. Department of Environmental and Information Studies; Tokyo City University; 3-3-1 Ushikubo-nishi, Tsuzuki-ku Yokohama 224-0015 Japan
6. Graduate School of Media and Governance; Keio University; SFC Fujisawa Kanagawa 252-0882 Japan
7. Biology Department; Boston University; 5 Cummington Street Boston MA 02215 USA
8. Department of Ecology and Evolutionary Biology; Yale University; 165 Prospect Street New Haven CT 06520 USA