1. Ministry of Education Key Laboratory of Southwest China Wildlife Resources Conservation, College of Life Sciences China West Normal University Nanchong Sichuan China
2. College of Life Sciences Sichuan Normal University Chengdu Sichuan China
3. Laojunshan National Nature Reserve Administration Pingshan Sichuan China
4. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, College of Life Sciences Beijing Normal University Beijing China