1. Department of Ecology and Evolution, University of Lausanne, Lausanne, Switzerland
2. Division of Aquatic Ecology, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland
3. Department of Fish Ecology and Evolution, Centre of Ecology, Evolution and Biogeochemistry (CEEB), Eawag Swiss Federal Institute of Aquatic Science and Technology, Kastanienbaum, Switzerland