1. Program in Ecology, Evolution, and Conservation Biology University of Illinois at Urbana‐Champaign Urbana Illinois U.S.A.
2. Illinois Natural History Survey, Prairie Research Institute University of Illinois at Urbana‐Champaign Champaign Illinois U.S.A.
3. Department of Natural Resources and Environmental Sciences University of Illinois at Urbana‐Champaign Urbana Illinois U.S.A.