1. New Zealand Institute for Advanced Study Allan Wilson Centre for Molecular Ecology & Evolution Massey University Private Bag 102904 Auckland 0745 New Zealand
2. Max Planck Institute for Evolutionary Biology August Thienemann Strasse 2 24306 Plön Germany
3. Laboratoire de Physique Statistique (UMR8550) École Normale Supérieure 24 rue Lhomond 75005 Paris France
4. Institut de Biologie de l'ENS (IBENS UMR 8197) École Normale Supérieure 46 rue d'Ulm 75230 Paris France
5. University Paris Diderot 75205 Paris CEDEX 13 France
6. Department of Ecology and Evolutionary Biology University of Arizona Tucson Arizona 85721
7. Laboratoire Ecologie‐Evolution UMR 7625 UPMC‐ENS‐CNRS École Normale Supérieure 46 rue d'Ulm 75005 Paris France
8. Institute of Natural and Mathematical Sciences Massey University Private Bag 102904 Auckland 0745 New Zealand