1. Department of Biological Science, Florida State University, Tallahassee Florida 32306
2. E‐mail:
3. Smithsonian Tropical Research Institute, Apartado 2072, Balboa, Republic of Panama
4. Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013‐7012
5. Center for Marine Biodiversity and Conservation, Scripps Institution of Oceanography, University of California San Diego, La Jolla, California 92093‐0202