1. School of Biotechnology and Biomolecular Science, University of New South Wales, Sydney 2052, Australia
2. Evolution & Ecology Center, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney 2052, Australia
3. E‐mail:
4. Now at Department of Biology, University of Minnesota, Duluth, 280 SSB, 1035 Kirby Drive Duluth, Minnesota 55812