1. EAWAG, Swiss Federal Institute of Aquatic Science and Technology, 8600 Dübendorf, Switzerland
2. ETH‐Zürich, Institute of Integrative Biology, 8092 Zürich, Switzerland
3. E‐mail:
4. Centre of Excellence in Evolutionary Research, Department of Biological and Environmental Science, PO Box 35, 40014 University of Jyväskylä, Finland
5. Faculty of Biological and Environmental Sciences, PO Box 56, 00014 University of Helsinki, Finland
6. Moskuantie 2 b 7, 90830 Haukipudas, Finland