1. School of Environmental and Evolutionary Biology; University of St. Andrews; St. Andrews Fife KY16 9TH United Kingdom
2. Laboratoire de phanérogamie; Institut de botanique de l'Université; Chantemerle 18, CH-2007 Neuchâtel Switzerland
3. NERC Centre for Population Biology; Imperial College at Silwood Park; Ascot Berkshire SL5 7PY United Kingdom
4. School of Biological Sciences; University of East Anglia; Norwich NR4 7TJ United Kingdom