1. Department of Biological Sciences; Graduate School of Science; The University of Tokyo; Hongo 7-3-1 Bunkyo-ku Tokyo 113-0033 Japan
2. Japan Science and Technology Agency; Precursory Research for Embryonic Science and Technology; 4-1-8 Honcho Kawaguchi Saitama 332-0012 Japan
3. Plant Global Education Project; Graduate School of Biological Sciences; Nara Institute of Science and Technology; 8916-5 Takayama Ikoma Nara 630-0192 Japan
4. Graduate School of Science and Technology; Kumamoto University; Kurokami 2-39-1 Kumamoto 860-8555 Japan