1. Guangdong Key Laboratory for Innovation Development and Utilization of Forest Plant Germplasm, College of Forestry and Landscape Architecture South China Agricultural University Guangzhou 510642 China
2. Biological Control of Pests Research Unit Agricultural Research Service, U.S. Department of Agriculture Stoneville MS 38776 USA
3. Department of Biology Salisbury University Salisbury MD 21801 USA
4. Department of Mathematics and Computer Science Salisbury University Salisbury MD 21801 USA
5. Beijing Key Laboratory for Forest Pest Control, College of Forestry Beijing Forestry University Beijing 100083 China