1. Univ. of Bordeaux, Integrative and Theoretical Ecology, LabEx COTE; Bât. B2 - Allée Geoffroy St-Hilaire FR-33615 Pessac France
2. CNRS, Inst. of Mathematics of Bordeaux; Talence France
3. Ifremer, LER Arcachon, Quai du Commandant Silhouette; Arcachon France
4. Irstea, Aquatic ecosystems and global changes Unit (UR EABX); Cestas France