1. Joint International Research Laboratory of Water and Nutrient in Crop, College of Resources and Environment Fujian Agriculture and Forestry University Fuzhou 350002 China
2. Fujian Provincial Key Laboratory of Plant Functional Biology, College of Life Sciences Fujian Agriculture and Forestry University Fuzhou 350002 China
3. Department of Biology, Hong Kong Baptist University and State Key Laboratory of Agrobiotechnology The Chinese University of Hong Kong Shatin Hong Kong 999077 China