1. Department of Geography and Geology; Hoffman Environmental Research Institute; Western Kentucky University; Bowling Green KY 42101 USA
2. Program in Sensory Physiology and Behavior; Marine Biological Laboratory; Woods Hole MA 02543 USA
3. Department of Ecology and Evolutionary Biology; Brown University; Providence RI 02912 USA
4. Marine Research Division; AZTI Tecnalia; Pasaia 20110 Spain