1. IPSiM Univ Montpellier, CNRS, INRAE Supagro 34060 Montpellier France
2. Umeå Plant Science Centre (UPSC), Department of Forest Genetics and Plant Physiology Swedish University of Agricultural Sciences 901 83 Umeå Sweden
3. UMR INRAE 1158 BioEcoAgro, BIOPI Biologie des Plantes et Innovation, SFR Condorcet FR CNRS 3417 Université de Picardie 80039 Amiens France