1. Hawkesbury Institute for the Environment Western Sydney University Penrith 2751 NSW Australia
2. Centre for Crop Health University of Southern Queensland Toowoomba 4350 Qld Australia
3. Institute of Ecology and Earth Sciences University of Tartu J. Liivi St. 2 504 09 Estonia
4. Department of Biological and Environmental Sciences University of Jyväskylä 40014 Jyväskylän yliopisto Finland