1. Erguna Forest‐Steppe Ecotone Research Station, CAS Key Laboratory of Forest Ecology and Silviculture, Institute of Applied Ecology, Chinese Academy of Sciences Shenyang China
2. College of Resources and Enviroment, University of Chinese Academy of Sciences Beijing China
3. Institute of Remote Sensing and Geographic Information System, School of Earth and Space Sciences, Peking University Beijing China
4. Institute of Ecology, College of Urban and Environmental Sciences, Peking University Beijing China
5. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, Coastal Ecosystems Research Station of the Yangtze River Estuary, Institute of Biodiversity Science, School of Life Sciences, Fudan University Shanghai China