1. Department of Biological Sciences Graduate School of Science Tokyo Metropolitan University Hachioji‐shi Tokyo Japan
2. Faculty of Bioindustry Tokyo University of Agriculture Abashiri Japan
3. Department of Agri‐production Sciences College of Agriculture Tamagawa University Machida Tokyo Japan
4. Systematic Botany Laboratory Department of Biological Sciences Graduate School of Science Tokyo Metropolitan University Hachioji‐shi Tokyo Japan
5. College of Science and Engineering Flinders University Adelaide SA Australia
6. Faculty of Agriculture Kagawa University Miki Japan