1. Marine Spatial Ecology Laboratory and Australian Research Council Centre of Excellence for Coral Reef Studies; School of Biological Sciences; The University of Queensland; St. Lucia Campus Brisbane QLD 4072 Australia
2. Marine Spatial Ecology Laboratory; Biosciences; College of Life and Environmental Sciences; Geoffrey Pope Building; University of Exeter; Stocker Road Exeter EX4 4QD UK
3. The Global Change Institute; The University of Queensland; St. Lucia Campus Brisbane QLD 4072 Australia