1. Department of Evolutionary Biology; Ecology and Environmental Sciences; and Biodiversity Research Institute (IRBio); Universitat de Barcelona; Avinguda Diagonal 643 08028 Barcelona Spain
2. TEREC unit; Department Biology; University Ghent; Ghent Belgium
3. Natural History Museum; Rotterdam The Netherlands
4. M. Smallegangehof 25; 4331 WC Middelburg The Netherlands