1. Department of Immunology; Faculty of Medicine; University of Tsukuba; Ibaraki Japan
2. Department of Immunology; Life Science Center of Tsukuba Advanced Research Alliance (TARA); University of Tsukuba; Ibaraki Japan
3. Department of Immunology; Japan Science and Technology Agency, Core Research for Evolutional Science and Technology (CREST); University of Tsukuba; Ibaraki Japan