1. Aquatic Ecosystems; Institute for Food and Agricultural Research and Technology (IRTA); St. Carles de la Ràpita 43540 Catalonia Spain
2. Laboratory of Marine Bioscience; Fukui Prefectural Univeristy; 1-1 Gauken-Cho Obama City Fukui 917-0003 Japan
3. Royal Botanic Garden Edinburgh; Edinburgh EH3 5LR United Kingdom