1. Department of Animal Science and Eco-tourism; College of Agriculture; Aksum University; PO. Box:314 Shire endasslasse Aksum Ethiopia
2. Wildlife Conservation Research Unit; Department of Zoology; University of Oxford; Tubney House Tubney OX13 5L U.K
3. Department of Animal; Rangeland & Wildlife Science; College of Agriculture; Mekelle University; Mekelle Ethiopia