1. School of Ocean and Earth Science University of Southampton National Oceanography Centre Southampton Waterfront Campus, European Way Southampton SO14 3ZH UK
2. Marine Biology and Ecology Research Centre School of Marine Science and Engineering University of Plymouth Drake Circus Plymouth PL4 8AA UK
3. Université du Québec à Rimouski Département de Biologie Chimie et Géographie 300 Allée des Ursulines Rimouski Quebec G5L 3A1 Canada