1. Department of Ecology and Evolutionary Biology University of California Santa Cruz CA 95064 USA
2. Wildlife Research Division Department of Environment Edmonton AB Canada
3. Departments of Anthropology and Biology The Pennsylvania State University 516 Carpenter Building University Park PA 16802 USA
4. Department of Biomolecular Engineering University of California Santa Cruz CA 95064 USA
5. Department of Bioinformatics and Genetics Swedish Museum of Natural History Stockholm SE‐104 05 Sweden
6. Department of Evolutionary Genetics Max Planck Institute for Evolutionary Anthropology Deutscher Platz 6 Leipzig D‐04103 Germany