1. CNRS; ISEM-UMR 5554; Montpellier France
2. University La Réunion/CRVOI; Ste Clotilde Réunion Island France
3. University Montpellier 2; Montpellier France
4. Institut Pasteur Tunis; Laboratory of Epidemiology and Veterinary Microbiology/; University Tunis El Manar; Tunis Tunisia
5. CNRS; CRBM-UMR 5237; Montpellier France