1. State Key Laboratory of Conservation and Utilization of Subtropical Agro‐bio‐resources and Guangxi Key Laboratory of Forest Ecology and Conservation College of Forestry Guangxi University Nanning Guangxi 530004 China
2. State Key Laboratory of Grassland Agro‐ecosystems School of Life Sciences Lanzhou University Lanzhou 730000 China
3. Institute of Ecology and Evolution University of Oregon Eugene OR 97403 USA
4. School of Biological Sciences University of Western Australia Perth WA 6009 Australia
5. Agriculture Botany Department Tanta University Tanta 72513 Egypt
6. Iriomote Station Tropical Biosphere Research Center University of the Ryukyus Uehara, Yaeyama Taketomi Okinawa 907‐1541 Japan
7. School of Environmental and Geographical Sciences University of Nottingham Malaysia Semenyih 43500 Malaysia