1. Department of Marine Biology; Tonekabon Branch; Islamic Azad University; Tonekabon Iran
2. Khorramshahr Marine and Technology University; Khorramshahr Iran
3. Department of Fisheries; Faculty of Agriculture Sciences and Natural Resources; Gorgan University; Gorgan Iran
4. Department of Fishery Science; Faculty of Natural Resources; Islamic Azad, University of Lahijan; P.O. Box 1616 Lahijan Iran
5. Department of Fisheries and Environmental Sciences; Faculty of Natural Resources; University of Tehran; PO Box 31585-431 Karaj Iran