1. Crop Genetics Department John Innes Centre Norwich Research Park Norwich NR4 7UH UK
2. Cell and Developmental Biology Department John Innes Centre Norwich Research Park Norwich NR4 7UH UK
3. National Key Laboratory of Plant Molecular Genetics CAS‐JIC Centre of Excellence for Plant and Microbial Science (CEPAMS) CAS Centre for Excellence in Molecular and Plant Sciences Institute of Plant Physiology and Ecology Chinese Academy of Sciences Shanghai 200032 China