1. Aquatic Ecology and Water Quality Management Group Wageningen University P.O. Box 476700 AAWageningen The Netherlands
2. Wageningen Environmental Research P.O. Box 476700 AAWageningen The Netherlands
3. Research Unit of Environmental and Evolutionary Biology, Namur Institute of Complex Systems, and Institute of Life, Earth, and the Environment University of Namur Rue de Bruxelles 615000Namur Belgium
4. Resource Ecology Group Wageningen University Droevendaalsesteeg 3a6708 PBWageningen The Netherlands
5. School of Life Sciences Nanjing University Nanjing210023 China
6. Institute of Ecology, College of Urban and Environmental Science, and Key Laboratory for Earth Surface Processes of the Ministry of Education Peking University 100871Beijing China