1. Eco&Sols, INRA, CIRAD, IRD, Montpellier SupAgro; Univ Montpellier; 34060 Montpellier France
2. Kellogg Biological Station; Michigan State University; 3700 E Gull Lake Dr Hickory Corners MI 49060 USA
3. Sorbonne Universités, UPMC Univ Paris 06, CNRS, IRD, INRA, Université Paris Diderot; Institute of Ecology and Environmental Sciences (UMR7618); 7 quai St Bernard 75005 Paris France